Every Q Branch scene compiled for your viewing pleasure.
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DESMOND LLEWELYN
Beginning with From Russia with Love, Desmond Llewelyn portrayed the character in every official film except Live and Let Die until his death in 1999. In the 1977 film The Spy Who Loved Me, as Q delivered the underwater Lotus, Major Anya Amasova/Agent XXX (Barbara Bach) greets Q as "Major Boothroyd".
Starting with Goldfinger, the notion that Bond and Q would have an often strained relationship with each other was introduced by Guy Hamilton; it continued in the series thereafter. While briefing Bond on the gadgets that he is going to use on his mission, Q often expresses irritation and impatience at Bond's short attention span, often telling him to "pay attention, 007", and Bond's playful lack of respect for his equipment, telling the agent, "I never joke about my work, 007". In Thunderball, Bond can be heard muttering "Oh no" when Q joins him in the Bahamas. A running gag appeared in later films where Q's prized gadget would be destroyed in a mishap often caused by necessity or Bond's recklessness – examples include the Glastron jet boat in Moonraker (Bond sends it over the Iguazu Falls to escape pursuit by Jaws), the Aston Martin Vantage in The Living Daylights (Bond is forced to prime its self-destruct device in order to evade the Czech police forces), and the BMW Z8 in The World Is Not Enough – which is cut in half by a helicopter buzz-saw.
However, on occasion, Q has shown a warm and fatherly concern for 007's welfare, such as at Bond's wedding in On Her Majesty's Secret Service, when he assures Bond that he is available if Bond ever requires his help. Q has also assisted Bond in a more active role in his missions in Octopussy, remaining to aid Bond in person even after another ally is killed, and Licence to Kill saw him travel to assist Bond while he is officially on leave from MI6 even after Bond has resigned from MI6 to pursue his own vendetta. He frequently refers to Bond as "007", rather than by his name. In GoldenEye, Q shares a joke with Bond for the first time, and in The World Is Not Enough when he reveals his plan to retire, Bond is saddened at the prospect.
This was the final film appearance of Desmond Llewelyn as Q in the James Bond series, although he would revive the role once again as Q in a Heineken commercial, a TV cross-promotion for The World Is Not Enough.
ALEC MCCOWEN
In the 1983 film Never Say Never Again, Bond received his gadgets from a man (played by Alec McCowen) he referred as Algernon and Algy. His opening line is "Nice to know old Q can still surprise you 00s," indicating that Q is an unseen character. In sharp contrast to the personality of Q in EON film series, Algy hopes to hear about "Lots of sex and violence" from James Bond following his mission. In the closing credits, he is named as "Q Algy". Q Branch itself is depicted as underfunded and ramshackle compared to the high-tech surroundings of the Eon films.
BEN WHISHAW
The character of Q did not appear in 2006's Casino Royale or its sequel, Quantum of Solace (2008). Bond actor Daniel Craig expressed concern over the character's absence, and expressed his hope that Q would return in Skyfall. In November 2011, it was announced that British actor Ben Whishaw had been cast in the role. In Skyfall, Q's gadgets were comparatively simple, consisting of a miniaturised radio and a gun coded to Bond's palmprint so only Bond could fire it. When Bond appears a little disappointed, Q comments 'Were you expecting an exploding pen? We don't really go in for that anymore,' in reference to a miniature grenade featured in GoldenEye. Q is demonstrated to be highly knowledgeable on the subject of computer security to the point where he designed some of the most sophisticated security protocols in existence. As with Desmond Llewelyn's Q, he also gets frustrated with Bond's knack for damaging or destroying the gadgets – at the end of Skyfall the Aston Martin DB5 is burned out in the final showdown with Silva.
In Spectre, Q assists Bond on his mission, similar to Q's assistance to Bond in 1989's Licence to Kill. Q meets Bond in Austria where he outruns SPECTRE agents after a ring he eventually decodes, revealing the organisation's existence. Q returns to London to assist Miss Moneypenny and M in foiling corrupt MI6 bureaucrat Max Denbeigh's launch of the Nine Eyes intelligence network. At the end he provides Bond with his remodelled Aston Martin DB5. He is the first Q who is younger than James Bond.